Mendelians and Biometricians 161
map as very rugged and difficult of ascent, not to be rushed by brilliant cavalry charges, but not impregnable before the persistent, slow, and methodical onslaught of a courageous and patient infantry; they are named the hills of "Masked Segregation." On the biometrical map they are marked as impregnable, when once occupied and entrenched, and are named "Continuous or Fluctuating Variations," or, in their more recent maps, as "Intermediates."
The great battle of the future is that which will be fought along this rugged range of the "Intermediates." The task of the Mendelian army is to take it. And, already in the plains below its brigades are beginning to deploy, and are making those initial dispositions which indicate that the assault is being prepared. At the same time, far away on the enemey's flank, in the valleys of Copenhagen, a great turning movement is being developed, and the brigades of the "pure lines" are preparing for their march along the dip-slope of the range, in order to strike the Biometrical army in its rear at the moment when the main Mendelian army unfolds its frontal attack up the rugged face of the escarpment.
Meanwhile, along the crest of the range, the concentration and entrenchment of the shaken centre of the Biometrical army is apparent. On the right, its wing which defended the village wherein the long tradition that evolution was almost wholly a matter of continuous variation, and that segregation of discontinuous variations played little part or none at all, has been hopelessly shattered. For everywhere that advancing and ardent left Mendelian wing has shown the evidence of such discontinuous variations and their segregation in the kingdoms of plants, animals, and Man. On its left, the Biometrical wing has been rolled back, and the position which defended the propositions that problems of inheritance can be solved by reference to one only of the two parents, that the characters of the offspring are determined by the summation in a regular series of ancestral increments, and that an advance knowledge of heredity can be gained by a indiscriminate massing together of zygotic characteristics, has been carried by
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